1 VIB Center for the Biology of Disease, Leuven, Belgium.
2 KULeuven Center for Human Genetics, Leuven Institute for Neurodegenerative Disorders (LIND), University Hospitals Leuven, and University of Leuven, O&N4 Herestraat, Leuven, Belgium.
3 Laboratory for Proteolytic Neuroscience, RIKEN Brain Science Institute, Wako-shi, Saitama, Japan.
4 Japan Science and Technology Agency, Saitama, Japan.
5 UCL Institute of Neurology, Queen Square, London, UK.